Name
Vachellia campechiana f. campechiana
Synonymy and types
Acacia cochliacantha Humb. & Bonpl. ex Willd., Sp. Pl. 4:1080. 1806. - Mimosa cohliacantha (Humb. & Bonpl. ex Willd.) Poir. in Lam. Dict. Suppl. 1:78. 1810. - TYPE: MEXICO. without exact locality [erroneously given as Guayaquil, Ecuador (Rudd 1966)], Humboldt & Bonpland s.n. (holotype: B, destroyed, F photo, M0 photo).
Acacia cymbacantha Zucc. ex Benth. Trans. Linn. Soc. London 30:501. 1875. - TYPE: MEXICO. W.Karwinski s.n. (holotype: M). NOTE: This name is listed in synonomy in the original description of Acacia cochliacantha with the note "may be the same species with very much larger spines, at least 1 in. long and ½ in. broad".
Poponax attenuata Britton & Rose, N. Amer. Fl. 23:90. 1928. - TYPE: MEXICO. PUEBLA: Río de Santa Lucia, in the vicinity of San Luis Tultitlanapa, near Oaxaca, Aug. 1908, C.A.Purpus 3050 (holotype: NY).
Formal description
Twigs, petioles, rachises and peduncles glabrous to lightly puberulent. Leaflet margins glabrous to lightly ciliate. Calyx lobes, corolla lobes, floral bracts and inflorescence receptacle glabrous to lightly puberulent.
Distribution
Disturbed relatively dry to moist sites from sea level to about 1800 m in western and southern Mexico.
Additional info
A common species of western and southern Mexico, Vachellia campechiana can be consistently separated from related species by the presence of flattened, spoon-shaped thorns that commonly exceed 30 mm in length and may be nearly as wide. On young twigs these thorns are not always developed, accounting for some misidentifications. Even on young twigs, however, the stipular thorns, though small, are usually flattened. In the field identification is much easier, as on older branches, and commonly on the main stems, enlarged spoon-shaped stipular thorns are common, though not always present on the young twigs. Even without the enlarged thorns, however, other characteristics may be used for consistent separation from V. pennatula and V. macracantha, its two closest relatives. The sessile petiolar glands, dense yellowish-green pubescences, and large number of leaflets per pinna in V. pennatula separate this taxon from V. campechiana. The mostly smaller leaves, leaflets, shorter pinnae length and fewer leaflets per pinna can be used to consistently separate V. campechiana from V. macracantha.
It is very likely that Vachellia campechiana hybridizes with V. macracantha. Rudd (1966) suggested that some specimens were difficult to place and considered it a strong possibility that hybridization was responsible. More recently Brimer et al. (1987) found that in areas of sympatry a few individuals were encountered that could not be conveniently assigned to either taxon. In a study of two populations, where both parents were common, individuals were encountered that fell outside the morphological range of these two taxa using Stepwise Discriminate Analysis (SDA). Most of the probable hybrid individuals clustered between the parental types on the SDA chart. In these individuals the leaflet size, and the distance between leaflets was typical of V. macracantha, while the shorter petioles and the flattened stipular thorns were typical of V. campechiana (Brimer et al. 1987).
Two forms of Vachellia campechiana have been recognoized, f. campechiana and f. houghii. These forms are sympatric throughout most of their ranges, though they may be ecologically or phenologically distinct. The only distinction between the two forms is the presence of a dense pubescence on the young twigs, petioles, rachises, and on the calyx and corolla lobes of f. houghii. Most accessions of Vachellia campechiana f. campechiana are not cyanogenic (Brimer 1984), however, Seigler and Ebinger (1988) found that 20% of 240 specimens examined tested positive for cyanogenic compounds. Of the 43 specimens of f. houghii tested for the presence of cyanide production, only five gave a moderate to strong positive test. The glycoside present is proacacipetalin, the same as that found in V. macracantha and a number of other acacias of this section (Seigler et al. 1978, Seigler and Conn 1982).
Key to the forms of Vachellia campechiana
a. Young twigs and petioles lightly pubescent to glabrous . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .Vachellia campechiana f. campechiana
a. Young twigs and petioles densely white lanate . . . .Vachellia campechiana f. houghii
Flowering time
April - July.
Representative specimens
MEXICO:
Baja California Sur:
- El Triunfo, 19 Jun 1981, D.S.Seigler & P.M.Richardson 11773 (ILL);
- 9.3 road miles S of turn-off to hwy. 19 along hwy. 1, S of La Paz, 17 Aug 1995, N.Snow & T.Prinzie 6517 (MEXU, MO);
Chiapas:
- a 2 km de Chicoasen, camino a Copainala, alt. 200 m, 13 Dec 1987, A.Reyes García, E.Martínez, G.Urquijo & A.Márquez 140 (MO);
- logging road from Rizo de Oro to Colonia Figueroa and Cerro Baúl, 23 May 1985, W.Thomas & J.L.Villaseñor 3620 (MICH);
Chihuahua:
- Mpio. Batopilas, N side of Barranca de Batopilas, alt. 990 m, 26 Feb 1973, R.A.Bye 3376 (CM);
- Mpio. Batopilas, E of La Bufa, alt. 3300 ft. 27-31 Oct 1974, R.A.Bye 5637 (ASU);
- Mpio. Batopilas, Barranca de Batopilas, vicinity of La Buff, 16 Jul 1974, R.A.Bye 5972 (CM);
- Mpio. Batopilas, E of La Bufa, alt. 3300 ft., 16 Jul 1974, R.A.Bye 6310 (CM, TEX);
- Santa Rosa, 31 May 1960, C.W.Pennington 287 (TEX);
- ca. 2 miles N from La Bufa on the road to Creel, 7 Jul 1969, O.H.Soule 565 (MO);
- ca. 2 miles N from La Bufa on the road to Creel, 7 Jul 1969, O.H.Soule 573 (MO);
- Barranca de la Bufa, 17 Sep 1983, R.Torres C. & P.Tenorio L. 3741 (MO);
Colima:
- W-side of bay across from Manzanillo, Jul 1956, D.P.Gregory & G.Eiten 312 (MICH, MO);
- 10 miles N of Colima, alt. 3700 ft., 6 Mar 1973, C.D.Johnson 370-73 (MO);
- 7 miles S of Colima, alt. 1100 ft., 7 Mar 1973, C.D.Johnson 382-73 (MO);
- 7 miles E of Armeria, 7 Mar 1973, C.D.Johnson 396-73 (MO);
- Estaciín Alzada, 26 Jun 1980, J.A.S.Magallánes 2325 (LL);
- a 5 km al NE de Los Ortices, 11 Sep 1980. J.A.S.Magallánes 2524 (MO);
- 16 miles WNW of Santiago, road to Cihuatlan, alt. 0-30 m, 22 Jun 1957, R.McVaugh 14967 (MICH, MO);
- steep bluffs above Río Salado, 5 miles S of Colima, alt. 400 m, 26 Nov 1959, R.McVaugh & W.N.Koelz 1084 (MICH, MO);
- 14 miles NE of Armeria on hwy. 110, 23 Jul 1975, D.S.Seigler & G.Holstein 9580 (ILL);
- 27 miles NE of Armeria on hwy. 110, 23 Jul 1975, D.S.Seigler & G.Holstein 9586 (ILL);
- 27 miles NE of Armeria on hwy. 110, 23 Jul 1975, D.S.Seigler & G.Holstein 9587 (ILL);
- 2 miles E of Armeria on hwy. 110, 23 May 1980, D.S.Seigler, P.M.Richardson & S.Thompson 11537 (ILL);
- along road from hwy. 15 to Playa del Oro, W of Santiago, 30 Aug 1973, W.D.Stevens & M.Fairhurst 1848 (MICH, TEX);
Durango:
- 10 miles N of Tamazula, alt. 2000 ft, H.S.Gentry 5265 (GH, MO, NY);
- Potrero San Juan, Rancho El Purgatorio, alt. 800 m, 22 Oct 1990, M.González, A.García & S.Acevedo 2562 (MICH);
Guerrero:
- Hornita Playa, about 2 miles E of Acapulco, 1 Jan 1944, F.A.Barkley 14052 (MO);
- km 145-146 on hwy. 51 from Tiquicheo to Zitacuaro, alt. 350 m, 3 Dec 1982, B.Benz, K.Benz & M.Burd 730 (MEXU, WIS);
- mountain side S of Taxco, 21 Jun 1935, O.M.Clark 7099 (MO);
- mountain side N of Acapulco, 24 Jun 1935, O.M.Clark 7222 (MO);
- road between Chilpancingo del los Bravos and Chichihualco, alt. 2260 m, 14 Jan 1979, T.B.Croat 45523 (MO);
- 6 km from Los Cedros, alt. 980 m, 28 May 1976, A.Delgado, M.A.Champomanes & O.Téllez 138 (MO) 4 miles SW of Gaucpato, alt. 1500 ft., 15 Jun 1954, H.A.Floyed & C.L.Ryan 109 (TEX);
- Chacamerito, Coyuca, 13 Jul 1934, G.B.Hinton 6286 (MO);
- Vallecitos-Ciruelos, 3 Jun 1937, G.B.Hinton 10265 (LL, MICH);
- Placeres, 8 Jun 1937, G.B.Hinton 10388 (LL, MICH, MO);
- 5.3 miles S of Chilpancingo on road to Chichihualco, 18 May 1983, W.C.Holmes 4431A (TEX);
- 17 km W of Teloloapán on hwy. 51, alt. 1700 m, 21-22 Nov 1971, H.H.Iltis & T.S.Cochrane 135 (WIS);
- 46 miles NW Ziluatanega, 7 Mar 1979, C.D.Johnson 668-79 (MO);
- near Acapulco, road to Pie de la Cuesta, alt. 4000 ft., 11 Jul 1941, W.Leavenworth & M.Leavenworth 943 (ILL, MO);
- Sierra Madre del Sur, N of Río Balsas, Temisco, alt. 220 m, 24 Nov 1937, Y.Mexía 8885 (MO);
- 12.2 miles E of Marquelia on hwy. 200, 12 Jun 1983, N.A.Murray & D.M.Johnson 1424 (MICH);
- cerca del Naranjo, 18 km al NE de Arcelia, alt. 600 m, 22 Mar 1979, J.C.Soto N. & S.Zarate 1269 (MO);
- 2 km al S de Tuzantlán, camino Inguala-Tuliman, 5 May 1982, D.Rodrígues & E.Martínez S. 2 (MO);
- 36 miles S of Iguala on hwy. 95, 27 May 1980, D.S.Seigler, P.M.Richardson & S.Thompson 11547 (ILL);
- 1 miles E of Juchitan, 28 May 1980, D.S.Seigler, P.M.Richardson & S.Thompson 11559 (ILL);
- Joya de el Zapote, alt. 1430 m, 24 Aug 1982, R.Torres C., P.Tenorio L. & C.Romero de T. 1126 (MO);
- Barranca del Limón, alt. 1350 m, 11 Mar 1983, Viveros J.L. & Casas A. 309 (MO);
Jalisco:
- 10-11 km al SWW de Tuxcacuesco, alt. 800 m, 24 Nov 1988, R. Cuevas & L. Gúzman 3365 (WIS);
- Mpio. Tuxcacuesco, 10-11 km al SW de Tuxcacuesco, alt. 800 m, 22 Feb 1990, R.Cuevas & L.Guzman 3697 (WIS);
- Trayecto entre El Higueral y Juan Gil Preciado, alt. 200 m, 16 Aug 1991, A.Flores M., N.P.Ramos G., & R.Omar B. 2941 (WIS);
- 17-19 km al SE de El Grullo, alt. 1000 m, 4 Jul 1990, L.Hernández & A.Vázquez 25 (CM);
- 2.5-3.5 km S of El Chaute along road to Manantlán, alt. 950-1050 m, 4 Jan 1979, H.H.Iltis, R.Gúzman M. &. M.Nee 1121 (WIS);
- Mezguitan, 4 miles S of 2nd bridge hwy. 80, alt. 1600 m, 29 Jul 1990, A.Jack & M.Sydor 064 (TEX);
- 15 miles W of Magdalena, 30 Dec 1972, C.D.Johnson 205-72 (MO);
- 3 miles S of Tecolatlán, alt. 4000 ft., 1 Jan 1973, C.D.Johnson 4-73 (MO);
- 15 miles W of Magdalena, alt. 1400 ft., 2 Mar 1973, C.D.Johnson 315-73 (MO);
- 5 miles S of Zacoalco, alt. 5100 ft., 6 Mar 1973, C.D.Johnson 349-73 (MO);
- 6 miles N of Atenquique, alt. 4000 ft., 6 Mar 1973, C.D.Johnson 351-73 (MO);
- 9 miles E of Cihuatlán, 8 Mar 1973. C.D.Johnson 420-73 (MO);
- 5 km al N de la Presa Cajon de Penas, 3 Jul 1980, J.A.S.Magallánes 2435 (MO);
- 3 km E of Nayarit State line on road from Guadalajara to Tepic, alt. 1300 m, 15 Jul 1969, B.Marcks & C.Marcks 1166 (WIS);
- 10-16 miles NE of Autlán, alt. 900-1400 m, 2 Oct 1960, R.McVaugh 19784 (MICH, MO);
- Estación de Biología de Chamela, 27 Apr 1976, L.A.Pérez & M.Pérez 1311 (MO);
- Mpio. Cuautitlán, 31.5 km al SSE de Autlán, alt. 700 m, 18 Dec 1987, M.Rosales & A.Flores 201 (WIS);
- 5 miles SW of Tecolotlán on hwy. 80, 21 Jul 1975, D.S.Seigler & G.Holstein 9546 (ILL);
- 45 miles NE of Guadalajara, alt. 3600 ft., 14 Sep 1960, B.C.Templeton 8813 (F, MICH, MO);
- Mpio. Techaluta, cerro al W de cruscero a Teocuitatlán, alt. 1350 m, 26 Jul 1994, A.Valencia, M.Huerta & J.Uribe 37 (WIS);
- 4-5 km al SW de Pedro Toxín, Tolimán Mpio., alt. 1000 m, 29 Jan 1987, A.Vázquez & L.Gúzmán 4106 (WIS);
- Mpio Autlán, W de Ahuacapan, alt. 950-1000 m, 27 Feb 1987, A.Vázquez & L.Guzman 4184 (WIS);
- Mpio Cuautitlá. 19 km al SE de Autlán, alt. 1900 m, 16 Jul 1987, A.Vázquez & M.Rosales 4310 (WIS);
México:
- Palmar, Dist. Temascaltepec, G.B.Hinton 4124 (NY);
- Vigas, Dist. Temascaltepec, 23 May 1935, G.B.Hinton 7418 (GH);
Michoacán:
- 7 km al N de Tiquichco, alt. 500 m, 12 Dec 1971, F.Chiang, G.Medrano & C.H.Ramos 504 (MO);
- 6 km de Agua de Obispo por carr. a Huétamo, alt. 420 m, 20 Jul 1983, H.M.Hernández & A.Chacon 176 (MICH, MO);
- Tacupa, District Huétamo, 14 Nov 1934, G.B.Hinton 6979 (MO);
- 1 mile S of Apatzingan, alt. 1200 ft, 5 Aug 1940, W.C.Leavenworth 477 (MICH, MO);
- a 2 km al W de San Jeronimo, camino a Los Chiles, alt. 290 m, 18 Jul 1982, E.Martínez & J.C.Soto N. 1439 (MO);
- 3 miles E of the border of Colima-Michoacán on hwy. 110, 23 May 1980, D.S.Seigler, P.M.Richardson & S.Thompson 11540 (ILL);
- a 6 km al SW de La Huacana, 31 Dec 1977, J.C.Soto N. 586 (MO);
- a 1 km al S de Paricuaro, 24 Dec 1977, J.C.Soto N. & R.L.Andrade 508 (MO);
- en El Infiernillo, municipio de Arteaga, alt. 150 m, 23 Dec 1981, J.C.Soto N. & A.R. de Soto 3684 (MO);
Morelos:
- cerro a 2 km al S de Quilamula, alt. 990 m, 5 Jul 1986, J.Bonilla 198 (TEX);
- 2 km al E de Quilamula, alt. 1200 m, 29 Aug 1991, J.Bonilla 1025 (MO);
- 2.5 km al N del Limón, alt. 1310 m, 29 Jul 1990, J.Bonilla & I.Lopez 964 (MO);
- 2 km al E de Quilamula, alt. 1200 m, 29 Aug 1991, J.Bonilla 1025 (MO);
- Trayecto Higueron-Tilzapotla, 21 Jun 1984, F.Fuchs 317 (MO);
- El Limón, alt. 1350 m, 19 Nov 1981, B.Guerrero C. 1348 (TEX);
- 3 km N of Alpuyeca on hwy. 95, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12609 (ILL);
- 11 km W of Jojutla, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12622 (ILL);
- 10 km from Jojutla on road to Valle de Vásquez, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12624 (ILL);
- 10 km from Jojutla on road to Valle de Vásquez, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12629 (ILL);
- 7 km W of Jojutla on road to Xaxocotla and Jojutla, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12635 (ILL);
- 7 km W of Jojutla on road to Xaxocotla and Jojutla, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12636 (ILL);
- 7 km W of Jojutla on road to Xaxocotla and Jojutla, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12637 (ILL);
- 7 km W of Jojutla on road to Xaxocotla and Jojutla, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12638 (ILL);
- 7 km W of Jojutla on road to Xaxocotla and Jojutla, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12639 (ILL);
- 7 km W of Jojutla on road to Xaxocotla and Jojutla, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12640 (ILL);
- 7 km W of Jojutla on road to Xaxocotla and Jojutla, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12641 (ILL);
- 7 km W of Jojutla on road to Xaxocotla and Jojutla, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12642 (ILL);
- 7 km W of Jojutla on road to Xaxocotla and Jojutla, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12643 (ILL);
- 7 km W of Jojutla on road to Xaxocotla and Jojutla, 19 Nov 1985, D.S.Seigler & J.E.Ebinger 12644 (ILL);
- 13 miles N of Amacuzas on hwy. 95, 27 May 1980, D.S.Seigler, P.M.Richardson & S.Thompson 11544 (ILL);
Nayarit:
- Margenes del Rio Santiago, 2 km rio arriba de El Ciruelo, alt. 360 m, 29 May 1992, R.Acevedo R. & J.Sosa L. 1279 (WIS);
- entre Villa Varadero y la desembocadura del rio Ameca, Bahía de Banderas, 11 Aug 1990, M.Cházaro B. & O.Montes 6321 (MO);
- Maria Madre, Tres Marias Islands, 23 Oct 1925, R.S.Ferris 5646 (MO);
- Mun. Nayar 12 km al W de la Mesa del Nayar, alt. 500 m, 11 Nov 1990, G.Flores F. & R.Ramírez R. 2421 (MO);
- 10 miles E of San Blas, 28 Feb 1973, C.D.Johnson 248-73 (MO);
- 18 miles SW of Compostela, 28 Feb 1973, C.D.Johnson 253-73 (MO);
- 2 miles SE of Las Varas, road to Mazatán, alt. 60-90 m, 12 Jul 1957, R.McVaugh 15352 (MICH, MO);
- Mun. Nayar Verda a 7 km de Jesús María, 5 Sep 1991, R.Ramírez & G.Flores F. 1024 (MO);
- Mun. Nayar 3 km al N del entronque a San Juan Peyotan, alt. 610, 31 Jul 1990, R.Ramírez R., G.Flores F. & O.Téllez V. 492 (MICH, MO);
- 4 km S of intersection to Acaponeta on hwy. 15, 25 Jun 1981, D.S.Seigler & P.M.Richardson 11799 (ILL);
- km 20-40 del camino de terraceria hacia la Presa de Aguamilpa, 15 Jan 1988, O.Téllez V. 11167 (MO);
- 8.5 km al E de el Tomellin, hacia Santa Maria, alt. 630 m, 5 Aug 1983, R.Torres C. & H.Hernández 3473 (MO);
Oaxaca:
- 5 km al NW de Las Margaritas, alt. 1050 m, 17 Jul 1987, R.Aguilar 266 (MO);
- 24.2 km al N de Laollaga, 10 Dec 1982, R.Cedillo T. & R.Torres C. 1982 (MO);
- 81 km SE Oaxaca, 21 Dec 1978, R.Conway & C.D.Johnson 297-78 (MO);
- 55 miles SE of Oaxaca on hwy. 190, alt. 1100 m, 2 Jul 1977, T.B.Croat 39943 (MO);
- Chinango, 29 Jul 1985, L.Hernández O. 81 (MO);
- gravel pit below S end of cortina, Temascal, 9 Jan 1964, D.H.Janzen s.n. (EIU);
- 4 miles N of Huajuapán, 7 Nov 1963, D.H.Janzen 375 (EIU, MICH);
- 2-4 km E of Tehuantepec, 1 Jul 1959, R.M.King 1175 (TEX);
- 13 km NE of Tehuantepec, 8 Jul 1959, R.M.King 1425 (TEX);
- beach of Laguna Superior, S of Juchitan, 11 Jul 1959, R.M.King 1569 (TEX);
- 9-10 km N of the village of La Ventosa, 15 Jul 1959, R.M.King 1669 (TEX);
- 2.5 km al N de Yosocuta en la carretera Huajuapán-Juxtlahuaca, 15 Jul 1976, A.S.Magallánes, R.Ramos & E.S.Magallánes 14 (MO, WIS);
- por el W de Rincón Bamba, al W de Salina Cruz, 12 Dec 1986, C.Martínes R. 802 (MO);
- Carnero 10 km al NW de Rincón Bamba, 8 Jul 1988, C.Martínez R. 1454 (MO);
- San Jose el Palmar, 17 Dec 1988, C.Martínes R. 2171 (MO);
- 5 km al E de Mariscala, alt. 1350 m, 3 Jul 1977, J.Rzedowski 34885 (MO);
- 50 miles E of Tehuantepec on hwy. 190, 3 Aug 1975, D.S.Seigler & G.Holstein 9779 (ILL);
- 3 miles N of Puerto Escondido, 29 May 1980, D.S.Seigler, P.M.Richardson & S.Thompson 11568 (ILL);
- Rancho del Santo, a 7 km al SW de Mancuernas, alt. 170 m, 20 Aug 1976, M.Sousa, C.Ramos, O.Téllez & L.Rico 5867 (MICH, MO, WIS);
- Aguazarea, 6 miles E of Río Grande, alt. 20 m, 5 Feb 1977, M.Sousa, O.Téllez, M.Ladd & J.C.Soto 7083 (MO);
- a 4 km del ojo de agua de Tlacotepec hacia la Ollaga, alt. 260 m, 26 Sep 1984, R.Torres C. & E.Cabrera 6272 (MO);
- Entomatado 20.5 km al SE de Puerto Escondido, 27 Apr 1983, R.Torres C. & R.Cedillo T. 2774 (MO);
- "La Junta" 8 km al SW de Huajuapán de León, 3 Apr 1982, R.Torres C. & P.Tenorio L. 177 (ILL, MO);
Puebla:
- 9.6 miles N of Puebla/Oaxaca border, km 45-46, hwy. 131, alt. 3800 ft., 28 Jan 1981. J.Bauml & M.Kimnach 366 (US);
- 5 miles S of Matamoros, near crest called Terpene, 2 Jul 1977, J.D.Dwyer 14296 (ILL, MO);
- Alrededores de Acatlán, 4 Feb 1978, R.Espino 19 (MO, UC);
- 10 km S de Acatlán (Río Acatlán), 6 Feb 1978, R.Espino 20 (MO, UC);
- Cantera Tlayua, 2 km al E de Tepeji de Rodríguez, alt. 1500 m, 19 Jan 1985, R.S.Felger 85-1 (MO);
- 5 km al SW de Santa Catarina Tehuixtla, alt. 1640 m, 22 Aug 1980, F.G.Medrano, V.Jaramill & J.L.Villaseñor F-1274 (MO);
- Tlanisloltepec, May-Jun 1909, C.R.Purpus 3867 (GH, MO, NY, US);
- en San Vicente Boquerón, 27 May 1985, J.C.Soto N. 8687 (MO);
- Mpio Jolalpan, Paraje Las Trojas a 5 km de El Salado, alt. 1200 m, 1 Sep 1985, J.M.Vidana Rodríquez 39 (ILL);
- 2 km NW of Tepexco on hwy. 160, 11 Dec 1990, D.Seigler, J.Ebinger, H.Clarke & C.Gratton 13043 (ILL);
- 18 km SE of Tepexco on hwy. 160, 11 Dec 1990, D.Seigler, J.Ebinger, H.Clarke & C.Gratton 13061 (ILL);
- on road from Izucar de Matamoros to Axachiapan, at road to Chietla, 27 May 1991, D.Seigler, J.Ebinger, H.Clarke & K.Readel 13567 (EIU, ILL);
- 28 miles NW of Izucar de Matamoros, 2 Aug 1975, D.S.Seigler & G.Holstein 9686 (ILL);
Sinaloa:
- 4 km al SW del Potrero, alt. 510, 8 Dec 1982, J.M.Aguilar, P.Tenorio & C.Romero 130 (ILL);
- Agua Caliente de Cebado, 26 Nov 1977, R.Espino 5 (ILL);
- N of Mazatlán, km 9 on side road to San Ignacio, 24 Oct 1978, J.E.Fryxell 154 (TEX);
- near Rosario, Senalva, no date, J.Gregg 1135a (MO);
- 8.2 miles NW of Rosario on hwy. 15, 9 Jun 1967, D.H.Janzen 1736 (EIU, ILL);
- 12 miles S of Guamuchil, 26 Dec 1972, C.D.Johnson 179-72 (MO);
- 33 miles S of Escuinapa, 6 Jan 1973, C.D.Johnson 120-73 (MO);
- 46 miles N of Mazatlán, 7 Jan 1973, C.D.Johnson 131-73 (MO);
- 38 miles S of Culiacan, 25 Feb 1973, C.D.Johnson 199-73 (MO);
- 38 miles S of Cubocan, 11 Mar 1973, C.D.Johnson 495-73 (MO);
- San Blas, 28 Jan 1927, M.E.Jones 23077 (MO);
- hwy. 15, Culiacan to Mazatlán at km 158 and 159, 27 Jun 1989, L.R.Landrum & S.S.Landrum 6289 (MO);
- foothills, La Noria, alt. 800 ft., 7 Oct 1925, Y.Mexía 126 (MO);
- Los Labrados, alt. 5 m, 14 Oct 1926, Y.Mexía 924 (F, GH, MICH, MO, NY, UC, US);
- 38 km N of Mazatlán on hwy. 15, 13 Jun 1981, D.S.Seigler & P.M.Richardson 11677 (ILL);
- 151 km S of Culiacan on hwy. 15, 13 Jun 1981, D.S.Seigler & P.M.Richardson 11680 (ILL);
- 118 km N of Culiacan on hwy. 15, 13 June 1981, D.S.Seigler & P.M.Richardson 11685 (ILL);
- N side of Topolobampo, 14 Jun 1981, D.S.Seigler & P.M.Richardson 11688 (ILL);
- 27 km N of Los Mochis on hwy. 15, 14 Jun 1981, D.S.Seigler & P.M.Richardson 11692 (ILL);
- 9 km N of Rosario on hwy. 15, 24 Jun 1981, D.S.Seigler & P.M.Richardson 11790 (ILL);
- 2 km N of Sinaloa-Nayarit border on hwy. 15, 25 Jun 1981, D.S.Seigler & P.M.Richardson 11800 (ILL);
- 2 km N of Sinaloa-Nayarit border on hwy. 15, 25 Jun 1981, D.S.Seigler & P.M.Richardson 11801 (ILL);
- 2 km N of Sinaloa-Nayarit border on hwy. 15, 25 Jun 1981, D.S.Seigler & P.M.Richardson 11802 (ILL);
- 2 km N of Sinaloa-Nayarit border on hwy. 15, 25 Jun 1981, D.S.Seigler & P.M.Richardson 11803 (ILL);
- 21 km S of Rosario on hwy. 15, 25 Jun 1981, D.S.Seigler & P.M.Richardson 11805 (ILL);
- 10 km NE of Villa Unión on hwy. 40, 25 Jun 1981, D.S.Seigler & P.M.Richardson 11806 (ILL);
- 31 km NE of Villa Unión on hwy. 40, 25 Jun 1981, D.S.Seigler & P.M.Richardson 11807 (ILL);
- 31 km NE of Villa Unión on hwy. 40, 25 Jun 1981, D.S.Seigler & P.M.Richardson 11808 (ILL);
- 31 km NE of Villa Unión on hwy. 40, 25 Jun 1981, D.S.Seigler & P.M.Richardson 11809 (ILL);
- Agua Caliente de Zevado, 8 Jan 1976, G.Shapiro 295 (LL);
- road to Guamuchil, 34 miles by road NW of Culiacan, alt. 400 ft., 3 Jul 1969, G.L.Webster & G.J.Breckon 15565 (GH, MO);
Sonora:
- El Mqagistral, 15 km al E de Concordia, alt. 230 m, 7 Dec 1882, J.M.Aguilar, P.Tenerio & C.Romero 118 (MO);
- 4 km al SW del Potrero, alt. 510 m, 8 Dec 1882, J.M.Aguilar, P.Tenerio & C.Romero 130 (MO);
- 6.4 miles by road (Mex. 15) N of Navajoa, 6 Aug 1960, R.S.Felger 3732 (ASU);
- San Bernardo, Río Mayo, 24 Aug 1935, H.S.Gentry 1618 (A, GH);
- hills behind Alamos, 16 Jul 1971, A.C.Gibson & L.C.Gibson 2065 (LL);
- Guaymas, 25 Jan 1927, M.E.Jones 23076 (GH, MO, NY, UC);
- Hermosillo, 20 Oct 1926, M.E.Jones 22492 (MO);
- slopes along el Río Bonito, 7 Oct 1939, C.H.Mueller 3694 (TEX);
- arroya de Cocoragua, alt. 60-70 m, 1 Aug 1935, F.W.Pennell 19486 (GH, US);
- cerca Arroyo El Chiltepin, 3.3 lm NNE Real del Alamito, alt. 270 m, 25 Feb 1996, A.L.Reina-G., T.R. Van Devender, L.C.Loyola-R. 96-13 (ASU);
- 127 km S of Cd. Obregon on hwy. 15, 14 Jun 1981, D.S.Seigler & P.M.Richardson 11697 (ILL);
- 1 km NW of Bahía San Carlos, 15 Jun 1981, D.S.Seigler & P.M.Richardson 11710 (ILL);
- Santa Ana, 25 km al W de San Nicolas, 28 Sep 1983, R.Torres C. P.Tenorio L. 3887 (MO);
- "Cerro Prieto", 15 km al NE de Navojoa, 2 Oct 1983, R.Torres C. P.Tenorio L. 3910 (MO);
- N of Banamichi where Hwy. 21 crosses Río Sonora, 17 Jul 1978, J.Vaughn 641 (MO);
- 1.5 km N of Arispe, 28 Jul 1978, J.Vaughn 675 (MO);
- 9 miles NW of Colorado, between Colorado and Mazatlán, 6 Sep 1941, I.L.Wiggins & R.C.Rollins 333 (MO);
Veracruz:
- alrededores de Cotaxtla, 28 Jun 1970, J.González G. 81 (MEXU);
- a 11 km de Actopan, alt. 250 m, 23 Nov 1970, A.Lot 1056 (MO);
- Tamarindo, alt. 200 m, 13 Nov 1970. J.Rzedowski 2848 (TEX);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12543 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12544 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12545 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12546 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12547 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12548 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12549 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12550 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12551 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12552 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12552 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12553 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12554 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12555 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12556 (ILL);
- edge of El Tamarindo on hwy. 140, 18 Nov 1985, D.S.Seigler & J.E.Ebinger 12557 (ILL);