Name
Vachellia rigidula (Bentham) Seigler & Ebinger, Phytologia 87: 166. 2005.syn. Acacia rigidula Bentham, London Jour. Bot. 1: 504. 1842.
Synonymy and types
Basionym: Acacia rigidula Benth., London J. Bot. 1: 504. 1842. Acaciopsis rigidula (Benth.) Britton & Rose, N. Amer. Fl. 23: 94. 1928. - TYPE: UNITED STATES. TEXAS: T. Drummond 161 (holotype: K).
Formal description
Shrub or small tree to 4 m tall. Bark light gray to brown, smooth to shallowly furrowed. Twigs dark purplish brown to dark gray, slightly flexuous, glabrous to lightly puberulent. Short shoots commonly present above the stipular spines, to 4 mm long, covered with acuminate stipules and old leaf bases. Leaves alternate, also commonly clustered on the short shoots, 2-9 mm long. Stipular spines light gray, symmetrical, terete, straight, thin, to 50(80) x 2.1 mm near the base, glabrous to lightly puberulent. Petiole adaxially grooved, 2-5(9) mm long, densely puberulent; petiolar gland solitary, mostly located medially on the petiole, sessile, circular, 0.3-0.6(1.1) mm across, apex depressed, glabrous to puberulent. Rachis absent. Pinnae 1 pair per leaf, 5-20 mm long. Petiolules 1-3 mm long. Leaflets (2)3 to 5 pairs per pinna, mostly opposite, 1.5-5.0 mm between leaflets, oblong to elliptic, terminal leaflets (4)6-13(16) x 3-6(8) mm, lateral leaflets slightly smaller, glabrous to rarely lightly pubescent beneath, lateral veins obvious, 3 to 5 veins from the base, base oblique, margins not ciliate, apex obtuse and usually mucronate. Inflorescence a loosely flowered elongated spike, 10-35 mm long, solitary or in clusters of 2 to 4(8) on the short shoots. Peduncles 0-3 x 0.4-0.7 mm, puberulent. Involucre usually 4-lobed, at the base of the peduncle, the lobe apex ciliate the remainder glabrous to puberulent, persistent. Floral bracts spatulate, 0.4-0.8 mm long, puberulent, deciduous. Flowers sessile, white to cream-colored; calyx 4-lobed, 0.4-0.8 mm long, puberulent; corolla 4-lobed, 1.2-1.7 mm long, glabrous or nearly so; stamen filaments 2.7-4.0 mm long, distinct; ovary glabrous to densely pubescent, on a stipe to 0.3 mm long. Legumes dark reddish brown, straight to slightly curved, flattened, constricted between the seeds, linear, 40-100 x 3-6 mm, coriaceous, reticulately striate, puberulent, eglandular, dehiscent; a chartaceous pericarpic strip lining each valve; stipe to 7 mm long; apex acuminate and sometimes beaked. Seeds uniseriate, no pulp, dark brown, oblong, strongly flattened, 5.5-7.5 x 3.5-4.5 mm, smooth; pleurogram oblong to U-shaped, 1.1-1.9 mm across. Flowers in February to July. Chromosome number: 2n = 26 (Turner 1959).
Distribution
Open areas, disturbed sites, on rocky slopes and bluffs, along fence rows and along arroyos, particularly at lower elevations from southern Texas (Isely 1973, Turner 1959) and northern Mexico in the states of Coahuila, Jalisco, Michoacán, Nuevo León, Querétaro, San Luis Potosí and Tamaulipas.
Additional info
Commonly forms extensive thickets and extensively collected, Vachellia rigidula is relatively consistent in its morphological characteristics. Much of the variation found has to do with the difference in size of the primary leaves (those developing on rapidly growing shoots), and the usually smaller leaves that develop from the short shoots. In most specimens, the terminal pair of leaflets are less than 11 mm long, but may rarely exceed 16 mm. Also, there are rarely only two pairs of leaflets on a pinna, but three to five pairs are common. Pubescence of the spines, twigs, petioles, and petiolar glands also varies. Generally, the spines, twigs and petiolar glands are glabrous to lightly puberulent, but the petioles may be densely puberulent. Glabrous individuals are rarely found. No variation, however, was found in the test for cyanogenic glycosides. Of the more than 100 specimens tested, all gave negative results, even when emulsin was used.
Plants in populations of this species from the area south of San Antonio, Texas (Atascosa Co.), occasionally harbor ants in the spines (Seigler and Conn 1982, Seigler et al. 1982). This does not appear to represent a highly coadaptive interaction such as exists in other, better known ant acacias, for the ant species is found on a number of unrelated plants. Neither the ant nor the Vachellia species appear to have undergone any significant modification.
Some confusion has existed as to the correct name for this taxon. Bentham (1842, 1875) and Standley (1922) used the name A. amentacea and listed A. rigidula as a synonym. Recently, Turner (1959) considered both A. amentacea (Central Mexico) and A. rigidula (Texas and adjacent Mexico) distinct species. More recently Isely (1969) suggested that A. rigidula and A. amentacea are conspecific. It is now apparent that the name A. amentacea has been incorrectly applied (see discussion under V. bilimekii).
Flowering time
February-July.
Representative specimens
MEXICO:
Coahuila:
- 15 miles S of Baranora, alt. 2800 ft., 16 May 1957, H.S.Gentry 16625 (LL);
- Nueva Posita, 25 Apr 1965, F.W.Gould 11240 (TEX);
- 64 km al NW de Múzquiz, alt. 500 m, 21 Apr 1977, R.Grether 625 (LL);
- 35 miles S of Monclova, alt. 4700 ft., 2 Aug 1973, J.Henrickson 11737 (LL);
- 2 miles NW of Ocampo in the area called La Mota, 19 Mar 1974, R.Jansen 90 (WIS);
- 17 km SE of San Paulo on road from La Favorita, 21 Mar 1973, M.C.Johnston, T.L.Wendt & F.Chiang 10327 (MO);
- 20 miles NW of Melchor Múzquiz, 4 May 1967, D.L.Latorre 135 (TEX);
- 24 miles N of Nava on hwy. 57, 23 May 1983, D.Seigler, J.Kramer & E.Carreira 11975 (ILL);
- 19 miles S of El Sauz on hwy. 57, 31 May 1983, D.Seigler, J.Kramer & E.Carreira 12041 (ILL);
- 19 miles S of El Sauz on hwy. 57, 31 May 1983, D.Seigler, J.Kramer & E.Carreira 12042 (ILL);
- 19 miles S of El Sauz on hwy. 57, 31 May 1983, D.Seigler, J.Kramer & E.Carreira 12043 (ILL);
- 19 miles S of El Sauz on hwy. 57, 31 May 1983, D.Seigler, J.Kramer & E.Carreira 12044 (ILL);
- 19 miles S of El Sauz on hwy. 57, 31 May 1983, D.Seigler, J.Kramer & E.Carreira 12045 (ILL);
- 19 miles S of El Sauz on hwy. 57, 31 May 1983, D.Seigler, J.Kramer & E.Carreira 12046 (ILL);
- 19 miles S of El Sauz on hwy. 57, 31 May 1983, D.Seigler, J.Kramer & E.Carreira 12047 (ILL);
- 19 miles S of El Sauz on hwy. 57, 31 May 1983, D.Seigler, J.Kramer & E.Carreira 12048 (ILL);
- 19 miles S of El Sauz on hwy. 57, 31 May 1983, D.Seigler, J.Kramer & E.Carreira 12049 (ILL);
- 19 miles S of El Sauz on hwy. 57, 31 May 1983, D.Seigler, J.Kramer & E.Carreira 12050 (ILL);
- 22 miles S of the jct. 57 and 53, S of Monclova on hwy. 57, 18 May 1980, D.S.Seigler, P.M.Richardson & S.Thompson 11503 (ILL);
- ca. 82 miles N of Saltillo near Santa Elena, 26 Mar 1967, J.S.Wilson 12621 (TEX);
- San Lázaro, 16 Jun 1936, F.L.Wynd & C.H.Mueller 134 (MO);
Jalisco:
- Laguna de Ocote, 21 Mar 1987, B.T.Styles s.n. (MEXU);
Michoacán:
- Huétamo, San Lucas, 27 Feb 1934, G.B.Hinton 5698 (A);
- Mal Paso, Huétamo, 24 Jun 1935, G.B.Hinton 7274 (A);
Nuevo León:
- hillsides around Monterrey, 23 Feb 1936, J.T.Buchholz s.n. (ILL);
- La Avimeza, 16 Mar 1978, B.Calvert 3 (TEX);
- 80 miles SE of Nuevo Laredo, 12 Apr 1964, P.R.Castano 35 (TEX);
- 8 miles NE of Monterrey, 24 Apr 1960, J.Crutchfield & M.C.Johnston 5304 (TEX);
- 3 miles NE of Cerralvo, 24 Apr 1960, J.Crutchfield & M.C.Johnston 5315 (TEX);
- roadside just W of Leona, 29 Apr 1949, R.T.Clausen 7623 (GH);
- Monterrey, 1891, C.K.Dodge 81 (MO);
- below Ebanito, alt. 740 m, 12 Mar 1980, G.B.Hinton 17731 (TEX);
- Obispado, alt. 500 m, Feb 1947, M.M.Lucas 202 (F);
- Huasteca Canyon, 10 miles W of Monterrey, 8 Jul 1953, W.E.Manning & M.S.Manning 53119 (GH);
- Sabinas Hidalgo, N of Monterrey, 7 Jul 1953, W.E.Manning & M.S.Manning 53112 (GH);
- 6 miles W of Ciudad Victoria , road to Jaumave, 21 Jul 1953, W.E.Manning & M.S.Manning 53393 (GH);
- along hwy. 85 near La Gloria, 3 Jan 1983, D.Neill 5310 (MO);
- Sierra Gomas, Bustamante Canyon, 12 Aug 1988, T.F.Patterson 6482 (TEX);
- Cañon de Huajuco, 20 Aug 1946, E.A.Pequeña, J.B.Paxon & F.A.Barkley 16M632 (MO);
- Stark's farm, Montemorelos, W.M.Pichon 267 (EIU);
- near Monterrey, 11 Jun 1889, C.G.Pringle 2526 (CM, LL, MO);
- Hacienda of El Carrizo, alt. 1200 ft., 28 Feb 1906, C.G.Pringle 13724 (MO);
- Cañon del Diente, 15 Jul 1970, L.G.Reinhard 8 (EIU);
- W side of hwy. to Miguel Alemán City, 3 Aug 1975, J.F.Rodríguez 24 (WIS);
- 11 km NW of Allende on Mexico 85, 17 Dec 1990, D.Seigler, J.Ebinger, H.Clarke & C.Gratton 13133e (EIU, ILL);
- W side of Cerro de la Silla, Monterrey, 18 Dec 1990, D.Seigler, J.Ebinger, H.Clarke & C.Gratton 13156 (ILL);
- S of Cadereyta near Monterrey, 19 Dec 1990, D.Seigler, J.Ebinger, H.Clarke & C.Gratton 13176e (EIU, ILL);
- S of Cadereyta near Monterrey, 19 Dec 1990, D.Seigler, J.Ebinger, H.Clarke & C.Gratton 13179e (EIU, ILL);
- 2 miles N of China on route 40, 5 Jun 1991, D.Seigler, J.Ebinger, H.Clarke & K.Readel 13755 (ILL);
- E side of Huasteca Canyon, 16 Jul 1975, D.S.Seigler & G.Holstein 9335 (ILL);
- 10 miles N of Montmorelos on hwy. 85, 7 Jun 1983, D.Seigler, J.Kramer & E.Carreira 12125 (ILL);
- a 5 km de La Escondida, por la carretera hacia Aramberri, alt. 1500 m, 21 May 1982, R.S.Silva 46 (ILL);
- Rancho Resendez, 26 Jun 1937, M.Taylor 379 (MO);
- 3 miles SW of Monterrey, 1 Jul 1956, B.L.Turner 3973 (TEX);
Querétaro:
- 11 km al NW de Jalpán, alt. 650 m, 18 Mar 1985, R.Fernández N. 2774 (NY);
- 2 km al NW de el Rancho Las Flores, cerca de Tancoyol, alt. 600 m, 3 Apr 1987, R.Fernández N. 3803 (NY);
- main road running NE from Jalpán de la Sierra toward Río Verde, about 49 km from Jalpán, alt. 1000 m, 10 Feb 1992, C.E.Hughes, S.A.Hughes & R.Atkinson 1602 (MO, NY);
- 7 miles NW of Jalpán on road to Río Verde, 3 June 1991, D.Seigler, J.Ebinger, H.Clarke & K.Readel 13679 (ILL) Cañon del Río Estórax, alt. 650 m, 5-6 Mar 1996, S.Zamudo 9670 (TEX);
San Luís Potosí:
- 4 miles E of Valles on Tampico hwy., alt. 700 ft., 14 Feb 1960, J.Crutchfield & M.C.Johnston 5125 (TEX);
- limestone hills, Las Palmas, alt. 300 ft., 7 Mar 1899, C.G.Pringle 8008 (CM, MO);
- 13 miles E of Río Verde on Mexico route 70, 18 May 1991, D.Seigler, J.Ebinger, H.Clarke & K.Readel 13477 (EIU, ILL);
- 15 miles E of Río Verde hwy. 70, 5 Jun 1983, D.S.Seigler, J.Kramer & E.Carreira 12078 (ILL);
- 8 miles E of Tamuín on hwy. 70, 6 Jun 1983, D.S.Seigler, J.Kramer & E.Carreira 12097 (ILL);
- 10 miles N of Cd. Valles on hwy. 85, 9 Jun 1980, D.S.Seigler, P.M.Richardson & S.Thompson 11626 (ILL);
- 14 miles W of Río Verde on route 70, 3 Jun 1991, D.Seigler, J.Ebinger, H.Clarke & K.Readel 13701 (ILL);
Tamaulipas:
- 2 miles N of Santa Teresa on hwy. 101, 22 Mar 1964, D.Ahshapanek 101 (TEX);
- 2 miles N of Santa Teresa on hwy. 101, 22 Mar 1964, D.Ahshapanek 108 (TEX);
- 2 miles N of Santa Teresa on hwy. 101, 22 Mar 1964, D.Ahshapanek 406 (TEX);
- between Ciudad Victoria and Matamoros on hwy. 101, 21 Mar 1965, D.Ahshapanek 495 (TEX);
- between Tamazunchale and Jácala, 23 Mar 1965, D.Ahshapanek 560 (TEX);
- on hwy. 85, 3 miles S of the edge of Ciudad Victoria, 30 Dec 1994, H.D.Clarke & K.Readel 96 (EIU, ILL);
- on hwy. 101, 10 miles NE of Ciudad Victoria, 1 Jan 1995, H.D.Clarke & K.Readel 111 (EIU, ILL);
- on hwy. 101, 10 miles NE of Ciudad Victoria, 1 Jan 1995, H.D.Clarke & K.Readel 113 (EIU, ILL);
- on hwy. 101, 10 miles NE of Ciudad Victoria, 1 Jan 1995, H.D.Clarke & K.Readel 114 (EIU, ILL);
- along hwy. 180-101 between Tampico and Matamoros, 16 miles S of San Fernando, alt. 100 m, 1 Mar 1987, T.B.Croat & D.P.Hannon 66015 (ILL);
- 15 miles E of Victoria, 4 Feb 1960, J.Crutchfield & M.C.Johnston 5022 (TEX);
- near San Filipe, alt. 200 ft., 6 Feb 1960, J.Crutchfield & M.C.Johnston 5043 (TEX);
- 1 mile W of Nuevo Morelos, alt. 1100 ft., 13 Feb 1960, J.Crutchfield & M.C.Johnston 5120 (TEX);
- 8 miles S of Villagrán, alt. 1600 ft., 12 Mar 1960, J.Crutchfield & M.C.Johnston 5241 (TEX);
- San Francisco, mpio. de Hidalgo, alt. 360 m, 21 Feb 1985, R.Diaz 278 (MO);
- Monterrey, 1891, C.K.Dodge 81 (MO);
- 16 miles S of Ciudad Victoria, 9 Apr 1961, R.M.King 4567 (TEX);
- 4-5 miles S of Ciudad Victoria, 18 Apr 1961, R.M.King 3823 (TEX);
- Morales, 15 Feb 1939, H.LeSueur 166 (TEX);
- vicinity of Victoria, alt. 320 m, 1 Feb-9 Apr 1907, E.Palmer 29 (MO);
- Ejido de San Antonio, km 132, carretera Victoria-Jaumave, alt. 1280 m, 5 Apr 1988, M.Martínez 293 (MO);vicinity of Victoria, alt. 320 m, 1 May-13 Jun 1907, E.Palmer 377 (MO, UC);
- 29 miles E of Cd. Mante on hwy. 80, 7 Jun 1983, D.S.Seigler, J.Kramer & E.Carreira 12112 (ILL);
- 15 miles S of Cd. Victoria on hwy. 85, 7 Jun 1983, D.S.Seigler, J.Kramer & E.Carreira 12121 (ILL);
- 1 mile S of Tres Palos on hwy. 101-180, 10 Jun 1980, D.S.Seigler, P.M.Richardson & S.Thompson 11633 (ILL);
- 4 km N of the San Luís Potosí border on route 85, 21 Dec 1990, D.Seigler, J.Ebinger, H.Clarke & Gratton 13217 (ILL);
Veracruz:
- 7 miles E of Ebano, 14 Feb 1960, J.Crutchfield & M.C.Johnston 5144 (TEX).
UNITED STATES:
Texas:
Atascosa Co.:
- sandstone hills, Campbellton, 9 Mar 1917, E.J.Palmer 11237 (MO);
- 9 miles S of Jourdantown, 18 May 1980, S.G.Saupe & J.Koebrich 187 (ILL);
- 9 miles S of Jourdantown, 18 May 1980, S.G.Saupe & J.Koebrich 188 (ILL);
- 9 mile S of Jourdantown on hwy. 16, 15 Sep 1979, D.S.Seigler & D.A.Young 11381 (ILL);
- 6 mile S of Jourdantown on hwy. 16, 20 May 1983, D.S.Seigler, J.Kramer & E.Carriera 11930 (ILL);
Bee Co.:
- between Peltus & Tuleta, 6 Mar 1954, M.C.Johnston 544 (TEX);
Brewster Co.:
- Big Bend National Park, near Solis, just N of Mariscal Canyon, 13 Sep 1961, D.S.Correll & M.C.Johnston 24565 (MO, TEX);
- Solis Ranch along Rio Grande, 22 Jun 1937, B.H.Warnock 580 (TEX);
- at mouth of Big Canyon, 24 Dec 1946, B.H.Warnock & L.C.Hinckley 461030 (TEX);
Calhoun Co.:
- 6 miles NE of Tivoli, 14 Mar 1971, W.R.LaForce s.n. (MO);
- ridges at Magnolia Beach, 14 Jun 1953, M.C.Johnston s.n. (TEX);
- Keller's Point, 9 Jul 1922, B.C.Tharp 1634 (TEX);
Cameron Co.:
- along Arroyo Colorado, 13 Mar 1964, D.S.Correll 28959 (LL);
- about 12 miles from Boca Chica, 17 Mar 1942, C.L.Lundell 10781 (LL, MO);
Dimmit Co.:
- 1 mile W of Carrizo Springs, 31 Mar 1959, M.C.Johnston 3834 (TEX);
- 3 miles NW of Carrizo Springs on US 277, 21 May 1983, D.S.Seigler, J.Kramer & E.Carreira 11965 (ILL);
- 25 miles from Eagle Pass on Carrigo Springs road, 29 Jul 1921, S.Ferris & C.D.Duncan 3033 (MO);
Duval Co.:
- ca 9 miles S of San Diego on hwy. 1329, 17 Mar 1969, W.F.Mahler 5278 (WIS);
- 1 mile S of intersection of farm road 2295 and hwy. 16, 20 May 1983, D.S.Seigler, J.Kramer & E.Carreira 11943 (ILL);
Edwards Co.:
- 48.6 miles S of Rocksprings, hwy. 674, 15 Apr 1968, J.A.Mears 2487 (TEX);
Frio Co.:
- 5 miles N of Pearsall, alt, 270 m, 14 Aug 1979, W.L.Wagner & J.C.Solomon 4087 (MO);
Goliad Co.:
- Goliad, 10 Mar 1916, E.J.Palmer 9132 (MO);
Hidalgo Co.:
- McAllen, 14 Feb 1937, C.E.R.Cameron 25 (TEX);
- E of Sullivan City, 1 Apr 1941, C.L.Lundell & A.A.Lundell 9880 (LL);
- Santa Ana National Wildlife Refuge, 14 Apr 1977, J.C.Solomon 2693 (CM);
- Santa Ana National Wildlife Refuge, 14 Apr 1977, J.C.Solomon 2739 (MO);
Jackson Co.:
- Vanderbilt, 10 May 1918, E.J.Palmer 9710 (MO);
- Vanderbilt, 10 May 1916, E.J.Palmer 9722 (MO);
Jim Hogg Co.:
- 9 miles S of Hebronville on hwy. 16, 20 May 1983, D.S.Seigler, J.Kramer & E.Carreira 11946 (ILL);
Jim Wells Co.:
- 29 miles NE of Freer on Farm Road 624, 10 Apr 1965, P.Malacara & F.Gutierrez, Jr. 56 (LL);
- gravel pit 9 miles N of Alice on route 281, 6 June 1991, D.Seigler, J.Ebinger, D.Clarke & K.Readel 13764 (EIU, ILL);
Karnes Co.:
- 3.5 miles N of Kenedy on hwy. 792, 25 Sep 1988, T.Davis 21 (WIS);
- immediately S of Karnes City, 9 Apr 1969, D.Isely 10633 (MO);
- 0.5 miles SE of Lent, 29 Jun 1952, J.C.Johnson 897 (TEX);
Kinney Co.:
- Anacacho Ranch, 1 Apr 1965, D.S.Correll & H.B.Correll 30773 (LL);
- 15 miles NE of Brackettville on ranch road 334, 20 May 1976, D.S.Seigler, S.G.Saupe & H.Welt 9931 (ILL);
LaSalle Co.:
- Encinal, 16 Apr 1965, F.Guajardo 56 (LL);
- 3 miles S of Dilley, 26 Feb 1944, J.T.Painter & F.A.Barkley 14309 (MO, TEX);
Live Oak Co.:
- 7 miles S of George West, 26 Dec 1946, C.C.Albers 46381 (TEX);
- 1 mile N of Swinney Switch, 28 Mar 1963, D.S.Correll 27060 (LL, MO);
- 4 miles NE of George West, 30 Mar 1967, R.G.Koch 2689 (WIS);
Maverick Co.:
- Eagle Pass, 10-14 May 1904, D.Griffiths 6346 (MO);
- Eagle Pass, 23 Mar 1932, M.E.Jones 29327 (MO);
- 2 miles N of Jct. of hwy. 57 and 277, 16 Sep 1979, D.S.Seigler & D.A.Young 11405 (ILL);
McMullen Co.:
- S of Tilden on Texas route 16, 11 May 1991, D.Seigler, J.Ebinger, H.Clarke & K.Readel 13267 (ILL);
- 4 miles N of Tilden on hwy. 16, 20 May 1983, D.S.Seigler, J.Kramer & E.Carreira 11933 (ILL);
- near Callahan, 24 Mar 1956, J.S.Thompson & B.L.Turner 26 (TEX);
Nueces Co.:
- Corpus Christi, alt. 40 ft., 5-12 Mar 1894, A.A.Heller 1382 (ILL, MO, TEX);
San Patricio Co.:
- Welder Wildlife Foundation about 7 miles N of Sinton, 26 Apr 1981, S.L.Hatch 4372 (TEX, WIS);
- 1 mile N of Sinton on US 77, 10 Jun 1980, D.S.Seigler, P.M.Richardson & S.Thompson 11635 (ILL);
- Welder Wildlife Foundation, N of Sinton, 5 May 1959, A.Traverse 1237 (MO);
- 3 miles NW of Mathis, 3 Apr 1960, G.C.Williges 369 (TEX);
Starr Co.:
- 8 miles N of Rio Grande City, 30 Aug 1932, E.U.Clover 472 (TEX);
- 3.5 miles S of La Gloria, 8 Feb 1969, D.S.Correll 36752 (LL);
- Falcon State Park, 19 Mar 1971, W.R.LaForce s.n. (MO);
- 2-3 miles SE of Sullivan City, 6 Apr 1978, J.Ortiz 19 (WIS);
- 3 miles N of Roma, 18 Apr 1965, J.W.Stanford 516 (TEX);
Uvalde Co.:
- Garner State Park, W of Uvalde, 18 Mar 1983, L.C.Barnett & L.J.Dorr 27 (TEX);
- 14 miles W of Uvalde, 19 Apr 1965, J.W.Stanford 526 (TEX);
- 15 miles N of Uvalde, 26 Jun 1954, B.L.Turner 3641 (TEX);
Val Verde Co.:
- 8.5 miles W of Comstock, route # 90, 3 Apr 1959, D.S.Correll 20718 (LL);
- 7 miles N of Del Rio on US 90, 27 Apr 1974, W.Hess 3170 (MOR);
- a little W of Langtry, alt. 1800 ft., 10 Mar 1950, L.C.Hinckley & L.Hinckley 409 (TEX);
- Seminole Canyon State Historical Park, 19 Mar 1988, Z.Labus 72 (TEX);
- Devils River N of Del Rio, 25 Apr 1903, H.A.Pilsbry s.n. (PH);
- 4.8 km W of Seminole Canyon State Park on hwy. 90, 18 Mar 1986, B.C.Reuter 314 (WIS);
- Rough Canyon camp ground, 23 Mar 1975, D.S.Seigler & T.L.Lockwood 8979 (ILL);
- Amistad Reservoir, 24 Jun 1974, D.Seigler, G.May & K.Hilu 8410 (ILL);
- Langtry, 21 May 1976, D.S.Seigler, S.G.Saupe & H.Welt 9955 (ILL);
- 10 miles W of Del Rio on US 90, 21 May 1976, D.S.Seigler, S.G.Saupe & H.Welt 9976 (ILL);
- Pecos River Bridge on US 290, 16 Sep 1979, D.S.Seigler & D.A.Young 11416 (ILL);
Webb Co.:
- Stony hills, Greene, 7 Apr 1901, H.Eggert s.n. (MO);
- 3 miles N of Laredo, 19 Mar 1964, J.M.Hughes 46 (LL);
- 30 miles N of Laredo on US hwy. 83, 2 Dec 1972, H.H.Iltis 27004 (WIS);
- 11 miles W of Encinal on route 44, 5 Jul 1960, H.H.Iltis, R.Koeppen & F.S.Iltis 44 (WIS);
- Lake Casa Blanca, 6 miles NE of Laredo, 8 Mar 1964, B.Notzon 55 (LL);
- 25 miles N of Laredo, hwy. 59, 28 Mar 1964, K.Peterson 124 (TEX);
- 4 miles N of Laredo on I-35, 21 May 1983, D.S.Seigler, J.Kramer & E.Carreira 11955 (ILL);
- 45 miles NE of Laredo on hwy. 59, 22 May 1976, D.S.Seigler, S.G.Saupe & H.Welt 10038 (ILL);
- 5 miles NE of Laredo on hwy. 59, 15 Sep 1979, D.S.Seigler & D.A.Young 11398 (ILL);
Willacy Co.:
- just S of Redfish Bay, 9 Mar 1954, M.C.Johnston 54190 (TEX);
Zapata Co.:
- 4 miles E of Zapata, Farm Road 496, 27 Mar 1964, L.Araiza, Jr. 49 (TEX);
- vicinity of Zapata, 7 Apr 1957, D.S.Correll & C.Schweinfurth 15723 (LL);
- 1 mile S of Falcon on US 83, 20 May 1983, D.S.Seigler, J.Kramer & E.Carreira 11954 (ILL);
- 2 miles N of San Ygnacio, 25 Mar 1964, M.E.Uribe 87 (TEX).