Key ex Holttum 1985, in litt.
| 1a. |
Veins all free |
T. dissecta |
|
Veins anastomosing |
2 |
| 2a. |
Veins forming regular costa areoles lacking free included veinlets; other areoles, if present, with few short, unbranched veinlets |
3 |
|
Veins copiously anastomosing with many, often branched, free veinlets in areoles |
11 |
| 3a. |
Upper surface bearing hairs between the veins |
4 |
|
Upper surface lacking hairs between veins, except near margins |
7 |
| 4a. |
Indusia absent of very small |
5 |
|
Indusia conspicuous |
6 |
| 5a. |
Stipe and rachis dark, glossy; axes or frond scaly on abaxial surface, not hairy |
"T. squamipes" |
|
Stipe and rachis not dark and glossy; axes of frond +/- hairy on abaxial surface, not scaly |
T. ferruginea |
| 6a. |
Additional areoles ocmmonly present below sinuses between lobes of pinnae or pinnules; hairs between veins on upper surface very thick |
T. pubescens |
|
Additional areoles below sinuses lacking or rare; hairs on upper surface slender; limestone plants |
T. devexa |
| 7a. |
Suprabasal pinnae with several pairs of free or adnate pinnules; sterile and fertile fronds isomorphous |
8 |
|
Suprabasal pinnae lacking free pinnules; fronds +/- dimorphous |
10 |
| 8a. |
Rachises dark and glossy, bearing numerous narrow scales on abaxial surfaces |
"T. nabirensis" |
|
Rachises lacking scales on aboxial surfaces |
9 |
| 9a. |
Basal acroscopic lobes of pinnules little longer than basal basiscopic ones |
"T. nesiotica" |
|
Basal acroscopic lobes of pinnules much longer than basal basiscopic ones |
T. kingii |
| 10a. |
Fronds opaque, smaller veins not visible on surfaces; buds present at bases of pinnae |
T. bamleriana |
|
Fronds not opaque; veins all distinct; no buds. |
T. brachiata |
| 11a. |
Lamina simple, its base cordate |
T. novoguineensis |
|
Lamina deeply 3 - 5-lobed or pinnate |
12 |
| 12a. |
Lamina deeply 3 - 5-lobed, its base cordate |
13 |
|
Lamina lobed to a winged rachis; stipe also winged |
15 |
| 12c. |
Lamina consisting of free pinnae, at least at its base; stipe unwinged |
17 |
| 13a. |
Sori indusiate, near margins; both surfaces copiously short-hairy |
T. craspedocarpa |
|
Sori exindusiate, irregularly scattered, sometimes coalescent |
14 |
| 14a. |
Lower surface of costae glabrous; rather many coalescent sori; basal scales short |
T. cristovalensis |
|
Lower surface of costae hairy; basal scales 15 mm or more long; fewer coalescent sori |
"T. croftii" (juvenille) |
| 15a. |
Sori in one row each side of main veins, +/- prominent on the upper surface; indusia large |
T. decurrens |
|
Sori exindusiate, not thus arranged |
16 |
| 16a. |
Sori rather regularly arranged in rows parallel to cross veins which join main veins; wing commonly 10-15 mm wide on each side of the rachis |
T. beccariana |
|
Sori very small, irregularly scattered; wing of rachis much narrower |
T. schultzei |
| 17a. |
Suprabasal pinnae entire or subentire |
18 |
|
Suprabasal pinnae deeply lobed |
30 |
| 18a. |
Sori in one row on each side of main veins |
19 |
|
Sori not in one row on each side of main veins |
26 |
| 19a. |
Indusium large and conspicuous; sori uniform |
20 |
|
Indusium small and thin or lacking; sori sometimes +/- enlongate or coalescing |
25 |
| 20a. |
At least basal pinnae auricled on their acroscopic side |
T. barclayi |
|
No pinnae auricled on the acroscopic side |
21 |
| 21a. |
Stipe and rachis dark and glossy |
22 |
|
Stipe and rachis not dark and gloosy |
23 |
| 22a. |
Bases of suprabasal pinnae broadly rounded; scales at base of stipe rigid, erect, dark twisted |
"T. exauriculata" |
|
Bases of suprabasal pinnae cuneate; scales at base of stipe otherwise |
"T. gymorachis" |
| 23a. |
Basal pinnae forked; indusia peltate or round-renform, receptacle not elongate along the vein |
24 |
|
Basal pinnae not forked; indusia +/- ellipsoid, receptacle elongate along the vein |
T. pleiosora |
| 24a. |
Apical lamina at most trilobed; lower surface of costae glabrous; indusia very firm, round reniform |
"T. isomorpha" |
|
Apical lamina commonly multilobed; lower surface of costae minutely hairy; indusium thinner, peltate to reniform |
T. crenata |
| 25a. |
Sori sometimes elongate or coalescent parallel to the main veins; fronds not dimorphous; pinnae to c.2 cm wide |
"T. refescens" |
|
Sori (especially the basal ones) elongate and sometimes coalescent at right angles to main veins; fronds usually dimorphous |
T. siifolia |
| 26a. |
Apical lamina trilobed, with 1 or more buds on the upper surface of its midrib; sori small, irregularly arranged, mostly on small veinlets in areoles |
T. angulata |
|
Apical lamina lacking buds; sori mostly not on free veinlets in areoles |
27 |
| 27a. |
Caudex creeping, to 6 mm diameter, basal pinnae unlobed, to c. 25 x 3cm |
T. menyanthidis |
|
Caudex erect or ascending, with tufted fronds; basal pinnae shorter, usually lobed |
28 |
| 28a. |
Fertile pinnae c.7 mm wide, much narrower than sterile, with simpler venation; sori nearly all coalescent, some coenosori linear and submarginal |
T. teratocarpa |
|
Fertile pinnae conform with sterile |
29 |
| 29a. |
Basal pinnae 4-5 cm wide above basal lobe; buds present at their bases |
"T. croftii" |
|
Basal pinnae to 2 cm wide above basal lobe; no buds at their bases |
"T. rheophytica" |
| 30a. |
Stipe and rachis dark |
31 |
|
Stipe and rachis not dark |
T. brachiata |
| 31a. |
Apical lamina many-lobed |
32 |
|
Apical lamina trilobed |
T. barclayi |
| 32a. |
Fronds dimorphous; areoles containing sori filling whole area of lobes of fertile pinnae |
T. durvillei |
|
Fronds not dimorphous; many areoles not containing sori; limestone plants |
T. melanorachis |
Key ex Copeland 1928, Philip. J. Sci. 37:403.
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