Medium size to very large terrestrial ferns. Rhizome fleshly, erect, massiveglobose, radially symmetic, vascular bundles in a complex polycyclic dictyostele, bearing +/- deciduous flat scales. Fronds erect, stipes fleshy, terete, with an enlarged base (pulvinus), subtended by 2 large stipule-like growths that may remain on the rhizome after the frond has fallen, lamina glabresent, thick and fleshy, bibinnate, +/- truncate, the basal and apical pinnae (an pinnules) only slightly reduced, pinnae pulvinate basally, pinnues free and shortly pulvinate, short-stalked, costate margins finely serrulate, at least apically; veins simple or forked, close and parallel, free to the margin, each vein or branch terminating in a tooth, recurrent false veins often present between each vein extending from the sinus. Sori in a single intramarginal row; sporangia large, thick-walled, free, along the veins in a short, close, double row, dehiscing by a longitudinal slit towards the vein; exindusiate, pluricellular uniseriate paraphyses usually present at the base of sporangia. Spores monolete, minutely pustular.